A) The fitness of a trait
B) The heritability of a trait
C) Whether a trait is an adaptation
D) Whether a trait is frequency-dependent
E) The level of selection on a trait

A) Determine the fitness of different trait values
B) Quantify the amount of variation in the trait in a population
C) Determine whether the mean trait value changes over time in a population
D) Conduct a parent-offspring correlation analysis of the trait
E) Determine whether the behavioral trait changes over time among individuals

A) Selection on groups does not occur
B) Top-down selection is stronger than bottom-up selection
C) Directional selection is stronger than stabilizing selection
D) Selection on groups may sometimes be stronger than selection on individuals
E) Kin selection is stronger than disruptive selection