What will NOT occur when 2,4-dinitrophenol (an uncoupler of oxidative phosphorylation) is added to actively respiring mitochondria?

A) The P:O ratio will increase.
B) The rate of NADH reoxidation at Complex I will increase.
C) The rate of oxidation of intermediates in the citric acid cycle will increase.
D) Proton export from the matrix will occur.
E) Heat will be released.