Which enzymes of the TCA cycle catalyze oxidative decarboxylation reactions?

A) malate dehydrogenase and citrate synthase
B) fumarase and succinate dehydrogenase
C) α-ketoglutarate dehydrogenase and succinate dehydrogenase
D) isocitrate dehydrogenase and α-ketoglutarate dehydrogenase
E) aconitase and succinate dehydrogenase