Nucleosomes that are positioned like beads on a string over a region of DNA can interact to form higher orders of chromatin structure. Which of the following factors can contribute to the formation of the 30-nm chromatin fiber from these nucleosomes?

A) Interactions that involve the histone tails of neighboring nucleosomes
B) Interaction of the linker histone H1 with each nucleosome
C) Binding of proteins to DNA or the histones
D) ATP-dependent function of chromatin remodeling complexes
E) All of the above