In maize,overall kernel color is controlled by both the endosperm and the aleurone colors.The Y allele produces a yellow endosperm while the homozygous recessive (y/y)state produces a white endosperm.Endosperm color is masked by the presence of a colored aleurone.The R and Pr genes control aleurone color.The combination of Pr and R alleles lead to a purple aleurone.The combination of pr and R alleles lead to a red aleurone.A homozygous recessive (r/r)produces a colorless aleurone,allowing the endosperm color to be expressed.Which allele is epistatic?