Who Formalized the Distinction Between Proximate and Ultimate Explanations of Behavior

Question

Who formalized the distinction between proximate and ultimate explanations of behavior?
A) E. L. Thorndike
B) Charles Darwin
C) George Romanes
D) C. Lloyd Morgan
E) Niko Tinbergen

Accepted Answer

Niko Tinbergen formalized the distinction between proximate and ultimate explanations of behavior, contributing significantly to the field of ethology and our understanding of animal behavior.