Which of the following statements best reflects the work of Kenneth Ross (1997) on fire ants?

A) He identified important gene-environment interactions that affect social behavior.
B) He used a knockout technique to understand variation in social organization.
C) He used QTL analysis to understand the ants' social behavior.
D) He used a linkage map to identify minor genes affecting the ants' behavior.
E) He identified a major gene that affected social organization.