Which principle is best supported by the results of Sutherland's studies in which fish predators were excluded from a marine fouling community?

A) Communities can follow different successional paths and display alternative states.
B) Predation is a relatively unimportant factor in succession in marine communities.
C) In most marine systems, the tolerance model accurately predicts successional outcomes.
D) Facilitation is most important in early succession, whereas inhibition becomes more important in later stages.