Which of the following is correct regarding the regulation and maintenance of Hox gene expression in Drosophila?

A) Proteins of the Polycomb and Trithorax groups maintain inactive and active states of Hox gene expression, respectively.
B) If Polycomb or Trithorax group genes are defective, Hox gene expression patterns are still initiated, but cannot be correctly maintained.
C) The remodeled chromatin at the Hox complex is heritable through cell generations.
D) If all the Hox genes in an embryo are deleted, segmentation still occurs but distinct segment identities are lost.
E) All of the above.