A giant aggrecan aggregate can have a molecular mass of 10⁸ daltons or more, and can be as large as a bacterium. Its structure resembles a brush. What is responsible for attaching the bristles of the brush to its core?

A) Covalent bonds between aggrecan core protein and keratan or chondroitin sulfate.
B) Noncovalent bonds between aggrecan core protein and keratan or chondroitin sulfate.
C) Covalent bonds between aggrecan core protein and hyaluronan.
D) Noncovalent bonds between aggrecan core protein and hyaluronan.